
A study of how a bird almost
became extinct in the 1930s, some 30 years before its
wetlands habitat started disappearing faster than it ever
had before, and 30 years before the human population started
growing rapidly. The irony is that the White-tailed Kite's
population became more stabilized during the time that the
human population was rapidly growing and the habitat was
disappearing. Over a thirty year period California
eliminated approximately 90% of its wetlands, and the
White-tailed Kite population grew.



12. Microtene
Bust
In 1960 the author was invited to go on a hastily planned field trip to an area just north of Marin County in northern California. A White-tailed Kite (Elanus leucurus) had been spotted and people were gathering to go take a look at this rare bird. For many it would be their first look. We eventually found the kite flying around a wet grassland looking for its favorite food, the California Vole (Microtus californicus).
The history of this bird in the United States and especially in California, is unique in the study of birds. In historical writings from 1886 through 1945, ornithologists were predicting the demise of this species. As Grinnell (1915) a California ornithologist, put it, the White-tailed Kite was due for "an early extinction."
Current records show that the kite's numbers have stabilized over the past 40 years. In 1927 Ralph Hoffman anticipated that there were only fifty pair of the birds in the entire state. Seventy years later there is good evidence that there are well over 2000 birds in California; from 100 birds to 2000 birds in seventy years.
How the White-tailed Kite managed to not become extinct and instead become a fairly common bird in its favorite habitat is a modern-day ecological mystery.
There are two goals of the White-tailed Kite Curriculum: (1) to develop a portrait of a species returning from the brink of extinction, and (2) to set up a curriculum that would introduce the student to using science to explore the dynamics of the White-tailed Kite.
|
The theme running through this book is that the structure of the environment, the morphology of the species, the economy of species behavior, and the dynamics of population changes are the four essential ingredients of all interesting biogeographical patterns. Robert MacArthur, Geographical Ecology - Patterns in the Distribution of Species |
The White-tailed Kite is a good opportunity for a study of biogreographical patterns. It is very connected to a specific type of habitat, the wetlands. The structure of the enviornment is important. The morphology, the structure of the Kite is well adapted to the wetlands habitat.
The White-tailed Kite is a specialist. A specialist is a species that has very specific requirments. A crow is not a specialist; it is a generalist. It can live in a variety of different habitats, eat whatever food is available, and can breed in a wide range of circumstances. A specialist is a species that has rigid requirements. An Osprey is a specialist. It eats only fish that it hunts from the water. The Snail Kite is a specialist as it eats only snails. The White-tailed Kite feeds primarily on voles, lives primarily in wetlands, and has a fairly specific range that it lives.
It is a medium-sized raptor whose morphology - wing structure, sharp talons, and accurate eye - combine to make it a skilled hunter. The kite is buoyant and quick and uses the wind to its advantage to catch its daily minimum of voles. It is found in grasslands lower than 2000 feet that are the results of fresh water supplies. It is so well adapted to its environment that it is one of few raptors that generally doesn't migrate. When the vole population explodes the kite can sometimes breed twice in the season to take full advantage of the increased number of voles.
The White-tailed Kite has gone from abundant to almost extinct to locally common over the past 100 years. As wetlands have disappeared and forest became farms, we have become aware of how the effect of modern civilization has been dangerous to our fellow animals. In the midst of these concerns it is intriguing to come across a species that was anticipated to become extinct and did not. The anticipation of its extinction happened even before its breeding grounds started disappearing in the later part of the twentieth century. Why did it not become extinct? What were the factors that enabled a bird that had practically disappeared to once again become a regularly breeding resident in a disapearing habitat?
The goal of The White-tailed Kite Curriculum is to create a science curriculum that explores some of the data involved in the history of this bird. Science is the study of the forces at work in our world. Generally the study of science does not utilize population studies of a bird as an approach to understand the broad truths of science.
Part One of The White Tailed Kite Curriculum introduces the natural history of the White-tailed Kite. It then provides an historical account of the status of this bird.
In order to document the status of this bird we will use numbers provided by the Christmas Bird Counts (CBC). CBCs have been conducted for over 100 years. By its longevity alone the information provided has become valuable data. Within California it is possible to study the density of kites in over 30 plots for over 30 years. The author is pleased to note that he participated in some of these counts.
In addition to the historical accounts and the CBC numbers, other chapters are utilized to introduce the science of population ecology and to compare the status of the White-tailed Kite and a very similar raptor, the Northern Harrier. What does the success of the Northern Harrier tell us about the White-tailed Kite? Can either species be considered a keystone species or an indicator species? How different is the Northern Harrier's habitat selection from the White-tailed Kite's habitat selection? What does the success of the Northern Harrier tell us about the White-tailed Kite?
From the USGS Glossary of Avian Conservation Terms - "Biogeography - the study of the geographic distribution of organisms, both past and present". We have 100 years of CBCs to develop a sketch of how the White-tailed Kite has survived over the past 100 years. The number of CBCs has grown at the same time that the population of the kite has grown. But we have good resources to show that the number of kites seen between 1900 and 1930 on the CBCs accurately represents the number of kites in California. It is a rare situation to have this kind of data that follows the rejuvenation of a species.
Part Two of the White-tailed Kite Curriculum takes the content from Part One and creates lesson plans for the independent student. The goal is to introduce a different approach to an ecological study.
|
|
|||||||||
|
|
|
|
|
||||||
The following sections are from a variety of different sources, that when combined form a general natural history of the White-tailed Kite. The natural history notes include six sections: Kites in general, breeding, food, hunting, habitat and distribution.
From Living Birds of the World :
|
The most primitive of the eight groups or subfamilies of diurnal birds of prey are the kites, which occur in most of the warmer regions of the world. They are long-winged and mostly long-tailed birds that are particularly adept at soaring, circling and gliding. They habitually work their way languidly over semi-open country, often in groups, their wings cocked at a slight dihedral angle. |
From, Birds of Prey of the World p. 207
|
The "black shoulder" is the common marking of this group, which is widely but erratically distributed throughout the warmer parts of the globe in steppe, desert, savanna, and the tropical forest. Four species are very similar, except for the slight variations in the underwing patterns. Three of them are considered by some ornithologists to be geographical representatives of a single species. |
White-tailed Kite (Elanus leucurus) formerly Black-shouldered Kite (Elanus caeruleus). Between 1961 and 1980 the White-tailed Kite was temporarily named the Black-shouldered Kite.
Breeding
Gillard, Living Birds of the World:
|
The White-tailed Kite (Elanus leucurus) about 16 inches in length, ranges from the southern borders of the United States through much of South America. Both sexes participate in nest construction, and four eggs are normally laid. Incubation requires up to 32 days, with both parents taking part. The young remain on the nest for about a month and then return to it for some time thereafter to obtain food. |
From, Birds of Prey of the World , p. 209
|
The White-tailed Kite builds a bulky nest of small fine twigs, deeply hollowed, and about 18 to 59 feet high in the top of an oak, willow, eucalyptus, or other deciduous tree. Both sexes take part in building, incubating eggs (about 30 days), and raising young, and the more aggressive individuals defend the nest. In southern California these kites group together in communities. They do not object to pairs of the same species building within 200 yards of their own nest, but the territorial boundaries are strictly observed. ... The normal egg clutch for Elanus is four, but varies from three to six. |
Food - Their diet is based largely on Microtine rodents. This is a large class of small rodents in the genus Microtus, generally known as voles. The most widespread vole in California is the California Vole, (Microtus californicus). Microtine rodents can have explosive population cycles. They can range from the density of 15,000 voles per hectare to 1 vole per hectare in just a few years. Frances Hamerstrom discusses a four year population cycle for the vole where in four years it goes from a high density of voles to an almost absence of voles.
From, Birds of Prey of the World p. 209
|
As in other species of Elanus, food includes small mammals (field mice, wood rats, pocket gophers, ground squirrels, shrews), reptiles, amphibians, and insects. They show a decided preference for living in the vicinity of fresh-water marshes and streams where food is readily available all through the year, and probably do not wander far, even in winter. |
From The Birder's Handbook; p. 222
|
Diet: Especially California vole and other rodents, also birds, snakes, lizards, frogs, and large insects. |
Hunting
From, Birds of Prey of the World p. 209 -
|
In the New World the White-tailed Kite often soars at a height of 60 or 70 feet while looking for prey on the ground. Remaining poised for half a minute at a time the bird then drops downward in an extended parachute maneuver to make the kill. Often it indulges in a pretty fluttering flight with quick wing beats. The display flight used in the nesting territory is a slow beating of the air with short strokes, the wings held at a sharp angle above the back, and the legs dangling. |
From Atlas of the Breeding Birds of Monterey County, California (1993) p. 88
|
The occurrence of the White-tailed Kite is closely related to the presence of voles and mice, and when the prey is depleted kites depart that area. Numbers and range seem to expand and contract in broad cycles, possibly tied to the abundance of its favored prey, the California Vole. For this reason the population is apparently cyclical in numbers, and breeding distribution may always be sporadic and unpredictable. |
From Shuford, David W. (1993). Marin County Breeding Bird Atlas. p. 133-135
|
Because Kites tend to be nomadic and to decrease or increase rapidly with fluctuating vole populations, future atlasers should be very cautious in interpreting any changes they detect when they repeat the Marin atlas at a later date. |
Habitat:
From California Birds - Their Status and Distribution
|
Open, cultivated and marshy bottomlands with scattered tall trees; savannah; grassy foothill slopes interspersed with oaks; agricultural areas with trees for windbreaks; orchards; and roadside verges (for hunting). They are found mainly in the Upper Sonoran Life Zone below 2000 feet, but wanderers occasionally occur in the mountains. |
From The Birder's Handbook p. 222
|
Breeding Habitat: Savanna, riparian woodland, marsh, partially cleared or cultivated fields, grassy foothills. |
From Grinnell - (1944) The Distribution of the Birds of California.
|
Habitat: Of two sorts: low rolling foothills or valley margins grown scatteringly to valley or live oaks; and river bottomlands or marshes adjacent to, or inclusive of, broken or scattering deciduous woodland. An essential combination of conditions seems to be open grasslands, meadows or marshes for foraging, primarily for meadow mice, and near-by isolated dense-topped trees for perching and nesting. |
Distribution:
From California Birds - Their Status and Distribution
|
Uncommon to locally fairly common resident. Although it is probably not truly migratory, the population scatters widely during the non-breeding season. Until recently winter roosts of more than 100 concentrated the population in localities from the Sacramento Valley and San Francisco Bay area south to San Diego County. Most out-of-range wanderers appear during fall, winter and spring. |
From, Birds of Prey of the World p. 209
|
The North American population has not been numerous in historic times and long ago disappeared from the southern part of the United States, except in Texas and California, west of the desert from the upper Sacramento Valley to the San Diego area. By 1900 the White-tailed Kite was considered rare in California and was protected by state law. Its beautifully marked eggs, some with a cap of brown coloring at one end, are prized by oologists. Part of the survival problem of this evidently dying species, like that of the Swallow-tailed Kite, has been interference by humans. |
Small, Arnold California Birds - Their Status and Distribution
|
Note: In recent years there had been an encouraging upsurge in the California population of these kites (which leveled off in the mid-1970s). This increase was probably due to an increase in the microtine rodent population as the result of new water projects, increased irrigation and the spread of agriculture. However, there was a steady and steep decline in coastal and near coastal southern California populations during the 1980s. Once the population leveled off, sharp local population fluctuations occurred, reflecting perhaps the cyclical situation in local rodent population. In some areas, the upswing in White-tailed Kites also indicated a reduction of local native floras and faunas due to conversion of indigenous vegetation to agriculture, and the expansion of conditions favoring both mice and kites. However, during the 1980s, the rapid urbanization of southern California and west-central California encroached upon agricultural lands, and may explain the movement of some kites to peripheral and marginal habitats (as into the fringes of the deserts), and the precipitous decline in areas where during the 1960s and 1970s they were fairly common. In some agricultural areas of the interior valleys, where kite populations were increasing during the 1960s and 1970s, conversion of crops favorable to microtine rodents resulted in unsuitable plantings of cotton, grapes and fruit orchards. Additionally, windbreaks, which served as nesting trees for the kites, were diminished. |
From The Birder's Handbook p. 222
|
Winters within US. North American range greatly expanded since 1960; probably only raptor to have benefited from agricultural expansion. Aided by high adaptability to habitat disruption and increased abundance of rodents. |
From Atlas of the Breeding Birds of Monterey County, California (1993) p. 88
|
Once considered endangered in California, the White-tailed Kite has made a great comeback in the state, being the only raptor to benefit from the spread of agriculture and the concurrent population explosion of its favored prey. |
Florence Merriam Bailey - (1902) Handbook of Birds of the Western United States
|
Distribution: Tropical America, except the West Indies; north in the United States to about the latitude of San Francisco on the Pacific coast, St. Louis in the interior, and South Carolina in the east. The kites are resident in the oak groves of Santa Clara Valley, and frequent the marshes about San Francisco Bay, where Mr. W. K. Fisher has found them catching large numbers of the California meadow mouse. |
Grinnell - (1915) A Distributional List of the Birds of California
|
It used to be particularly numerous around San Francisco Bay; but it is now everywhere very much reduced in numbers and restricted in range, with promise of an early extinction. |
Dawson - (1923) Birds of California, Volume 3, p. 1649
|
Recent ornithological literature bristles with records of occurances of the White-tailed Kite, once upon the verge of extinction, but, now, it is believed, being slowly nursed back into life. The cause of the near tragedy, now hopefully averted, was expressed several years ago by Dr. Grinnell: "With no doubt whatever, the present rarity of this hawk in California is due to the associational preference for marshes, where its habit of flying slowly back and forth at a moderate height above the ground in the lookout for meadow mice and insects makes it an easy target for the thoughtless hunter. In my experience the average sportsman is still unenlightened enough to shoot down any sort of 'hawk' that flies his way, provided game is not at the moment expected." Yet this is the bird of which Cooper (following Heermann) could write in the Sixties: "This beautful and harmless species is quite abundant in the middle districts of California, remaining in large numbers during winter among the extensive tule marshes of the Sacramento and other valleys." |
Hoffman - (1927) Birds of the Pacific States p. 64
|
There are probably not more than fifty pairs left in California and in spite of protection by law the number is slowly decreasing. |
Grosvenor, National Geographic Society's The Book of Birds - 1939
|
It is found over tree-dotted prairies and savannas, marshes, and semi-open valleys. Though fifty years ago it was common, it has decreased steadily until now it is to be classed among our unusual birds. Despite the fact that it has been afforded protection in recent years, the species does not seem able to increase. The White-tailed Kite is found in California from the upper Sacramento Valley and Humboldt County, south to northern Baja California and from Texas, Oklahoma, and Florida to Guatemala. |
Grinnell - (1944) The Distribution of the Birds of California.
|
Resident throughout year within metropolis of range; but shifts about locally, both seasonally and from year to year, in accordance with food supply. Formerly, prior to about 1895, common and widespread in valley and lower foothill territory, but now rare or entirely gone from many sections. A slight trend toward recovery, in area and numbers, is latterly in evidence. |
A. C. Bent , Life Histories of North American Birds
|
This gentle and attractive bird seems to have become exceedingly rare, or to have been entirely extirpated, in the eastern portions of its North American range. During my six seasons, or parts of seasons, spent in various portions of Florida I have never seen this kite; once a special trip was made to a section where our guide said they had recently nested, but no sign of them was found. Donald J. Nicholson tells me that he has not seen one there since 1910. We could not find it in southern Texas, and I have no recent records of it there. In certain sections of California it seems to be holding its own, though exceedingly local in its distribution, and nowhere universally abundant. I doubt if it ever was very abundant, although Cooper (1870) referred to it as "quite abundant in the middle districts of California, remaining in large numbers during winter among the extensive tule marshes of the Sacramento and other valleys", and Belding (1890) considered it "still a common resident" about these marshes "in the centre of the State." But Belding quotes Dr. B. W. Evermann, as calling it "a rare resident" in Ventura County, as early as 1886; and he quotes W. E. Bryant as saying that "it is still a very rare resident" in Alameda County. It seemed to be the general opinion, at that time, that the white-tailed kite was a disappearing species. As a result, it has since been rigidly protected by law and exempted from collecting permits. "Now comes more recent light on the subject, which is more encouraging. Dr. Gayle B. Pickwell (1930) has published the results of his exhaustive study of the literature and his field work in the Santa Clara Valley. Referring to past and present conditions in that region, he says: "In spite of the fact that Taylor, in 1889, wrote of the Kite, "I venture to assert that there are not more than four pairs this year breeding within a radius of seven miles of that city (San Jose)", today, forty-one years later, there are still that many or more. Let us estimate that an average of four pairs of Kites (too high an estimate for some, too low, perhaps, for others) frequents each. We have then sixteen pairs of Kites in this entire valley. Twenty pairs, forty birds, I feel convinced, account for every Kite from Gilroy to the Bay and from Mount Hamilton to the summit of the Santa Cruz Mountains. The Kite was certainly more numerous In San Joaquin and Sacramento counties forty to sixty years ago than it is now. In other regions where it was present, especially in marsh districts, undoubtedly It has been seriously reduced In numbers. The condition in hill sections inhabited by it can be but guessed at. Here it probably has suffered least. This Kite is probably a dying species, never within historical times having predominated as such raptorial birds as the Desert Sparrow Hawk or Red-tailed Hawk for instance. Since the above was written Dr. Pickwell (1932) has published a "requiem" for the kites in this valley; whereas he estimated that there were possibly 16 to 20 of these kites in the Santa Clara Valley in 1928, he now says: "This day (October 30, 1931) there cannot be more than two or three, and all too possibly none." We hope that this is a mere local condition. His observations on the home life of these kites were made in the foothills of the Mount Hamilton Range in Santa Clara County: The Slatore ranch lies in the foothills whose summits are grass-covered with wild oats and bromes, with scattered valley oaks and live oaks, and here and there a cluster of California coffee berry (Rhamnus catiforaica) and gnarled Sambucus. Rocky outcrops, where more moisture may be trapped, have curious copses of scrubby growths of toyon, holly-leaved cherry, sages and sage brush; and the gullies lined with buckeye, California laurel, and poison oak run down to Silver Creek where the laurels and willows predominate. But the hills are mostly smooth as velvet, golden velvet most of the year, and green oaks are scattered over the velvet, like buttons on a buxom vest. In three buttons on this velvet vest were occupied nests of the White-tailed Kite. That such a habitat is not an unusual Kite home is shown by the fact that all the Kites of Santa Clara Valley today are, excepting one or two pairs, restricted to the lower foothills of the Mount Hamilton Range and Santa Cruz Mountains, on either side of the north end of the Valley. The exception is of not more than two pairs that occur to the north of San Jose between that city and the Alviso salt marshes. These frequent the cottonwoods and eucalyptus trees of the Coyote Creek and, not infrequently, are seen hunting over the treeless marshes at the foot of the Bay in common with Marsh Hawks, native there, and Turkey Vultures and Red-tailed Hawks from the hills. Bendire (1892) says of their haunts: "Their usual resorts during the breeding season are the banks of streams or the fresh water marshes, especially if a few scattered live oaks or willow groves are close by, and their favorite nesting sites are the tops of live oaks, although other trees are also made use of whose foliage securely conceals the nest during incubation." The impression I gained from men I talked with in California and from my own limited experience there was that this kite shows a decided preference for the vicinity of water, fresh-water marshes and streams; in such places it finds its food readily available all through the year, and it probably does not wander far away even in winter. According to Audubon (1840) it was found in similar haunts in Texas and Florida. |
Ludlow Griscom Audubon's Birds of America (1950)
|
A rare and beautiful hawk now almost extinct in the Southeast, a few in Texas and California. Reapears in South America. |
From Atlas of the Breeding Birds of Monterey County, California (1993) p. 88
|
Historical Occurrence The overall distribution has dramatically changed over the past century. The increase in voles with the cutting of forests for agriculture has benefited this species. Once considered so rare that it was one of only five species fully protected under California law (along with the California Condor, Bald Eagle, Peregrine Falcon, and Trumpeter Swan; Fish & Game Code), the species has greatly expanded throughout the southern United States since 1960. In Monterey County kites were known from the Carmel Valley for years but the occurrence in the upper Salinas Valley is recent. No birds were recorded there by Willett (1908) but one bird was noted along the Nacimiento River on Camp Roberts by Mowbray (1947). Likewise, the spread to the Big Sur coast is recent, as the species was not seen by Jenkins (1906) or Pemberton & Carriger (1915). At Hastings Reservation, in the upper Carmel watershed, habitat is marginal for this species and kites occurred in only 17 of the 43 years between 1938-1980 and nested in only five of those years. |
From Shuford, David W. (1993). Marin County Breeding Bird Atlas. p. 133-135
|
Early declines were apparently caused by shooting, habitat loss, and perhaps by overzealous egg collectors. The eggs were highly prized because of their scarcity, because of the variability between egg sets, and because Black-shouldered Kite eggs are among the most beautiful of those of all North American birds. Illegal egg collecting continued until at least 1940 despite laws passed in 1905 to protect the birds and their nests (Williams 1940); further protection was afforded by legislation in 1957. California's Kite population has apparently increased because of the birds' ability to tolerate habitat fragmentation caused by agricultural practices, to exploit increased Microtus populations thriving in fields irrigated year round, and to reproduce at a high rate. A clutch size of four to five eggs and the ability to double brood in a single year are both unusual adaptations for a hawk ... Martin (1989) noted that the amount of irrigated agricultural land in California increased by 42% from 1944 to 1978, coinciding with the period of dramatically increasing Kite numbers. The great year-to-year fluctuations in Kite numbers appear, at least in part, to be tied to similar changes in the prey base influenced by rainfall. Pruett-Jones et al. (1980) found a significant positive correlation between Kite numbers and rainfall. This perhaps is explained by the fact that microtene rodents need standing water to reproduce ... and that their numbers are usually reduced in a drought. The decline in Kite numbers in California during the 1975-76 to 1976-77 drought and the substantial increase in numbers in Oregon at that time, including their first breeding record ... further suggests a link between rainfall, vole populations, and Kite populations. |
Christmas Bird Counts began in 1890 as a reaction by Chandler Robbins and friends to the wholesale slaughter of many of the nation's birds. Over the past 100 years private citizens have ventured out into their neighborhoods to conduct a census of wintering birds during the last two weeks of the year. In 2008 there were 2,111 Counts that totaled 57,700,668 birds.
A Christmas Bird Count is done by volunteers who break into teams and canvas a selected segment of a circle whose diameter is 15 miles long which equals 176 square miles and approximately 112,000 acres. The territory is a circle strategically drawn over a prime birding area. The circle is carefully segmented and teams are assigned. Some of the teams start before dusk and bird past 5 pm so they can hear owls and add them to the count record. Each count has a compiler and every compiler has a rare bird committee made up of seasoned experts to deal with occurrences of unusual birds. Efforts are made to keep the records as reliable as possible. The participants count every bird. If a flock of birds is seen then the number is estimated as best as possible. In the case of hawks and owls and other larger birds efforts are made not to count the same bird twice. Teams will communicate with adjacent teams to check notes to make sure that the bird didn't fly over into their territory.
I had a professor at San Jose State who told me that Christmas Bird Count data was worthless data. I don't agree with him. Perhaps one count by itself doesn't signify very much, but when you have over 100 years of CBCs (Christmas Bird Counts) and the citizen scientists who participate are as well skilled as they are, then the data is valuable; the trends significant.
The White-tailed Kite is easy to identify in the field. There are very few birds that can be confused with it. By its nature it stays out in the open where it can easily be seen. The problem is counting the bird twice, or three times as the bird wanders around to different areas that are being counted by different groups.
From the USGS Avian Conservation Glossary: "Index method - a counting method involving sampling that yields measures of relative abundance rather than density values." Is the CBC a determinant of relative abundance or density values? Density is determined for a circle but no effort is made to extrapolate to the whole area.
Landscape index: index of landscape structure (pattern), including richness, evenness, patchiness, diversity, dominance, contagion, edges, fractal dimension, nearest neighbor probability, and the size and distribution of patches.
|
|
|||||||||||
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
Table 2 NC = No
Count
Table 3
Auburn Los Banos 82 Rancho Santa Fe 15 Angwin La Purisima 11 Santa Ana River Valley 2 Ano Nuevo La Grange 17 Santa Barbara 43 Arcata Marin County 28 San Diego 17 Bakersfield Moss Landing 30 San Francisco 2 Benicia Milburn 2 Santa Maria 27 Bishop Mendocino 16 San Jose 19 Centerville Beach Monterey 3 South Fork Valley 1 Contra Costa County Moro Bay 34 San Jacinto Lake 40 Clear Lake Malibu 8 Sacramento 63 Claremont Oakland 11 San Bernardino 7 Chico Orange County 56 Springville 8 Carrizo Plains Orange County (Northeast) 29 Santa Rosa 41 Crystal Springs Oroville 8 Salton Sea South 5 Caswell Oceanside 31 Stockton 49 Escondido Palo Alto 26 San Juan Capistrano 14 Folsom Putah Creek 7 San Fernando 2 Hayward Pinnacles 1 Tule Lake 1 Hollister Parkfield 1 Thousand Oaks 32 Kern River Valley Palos Verdes 5 Ukiah 16 Los Angeles Point Reyes 43 Ventura 16 Long Beach Padadena 5 Willow Creek 1 Lake Henshaw Peace Valley 23 Western Sonoma County 7 Lost Lake Red Bluff 14 73 counts, 1425 Birds
identified Average per count = 20 Lancaster Redlands 3
The Northern Harrier is similar to the White-tailed Kite in many ways. They share very similar habitats. In practically every Christmas Bird Count that had a White-tailed Kite, there was also a Northern Harrier. There are more Northern Harriers than White-tailed Kite and they are found in more counts than White-tailed Kites. The harrier's diet is heavily dependent on microtine rodents. Both species hunt in very similar ways. While approximately 80% of the kite's population is found in California the Harrier's population is spread out through many states. The Northern Harrier has a migratory population in the states and a resident population that does not migrate. While the White-tailed Kite's diet is 99% California Voles, the Northern Harrier's diet is 56% California Voles.
The Red-tailed Hawk represents a raptor that is also a generalist and also has both a migratory and a resident population.
(Utilizing Christmas Bird Count Data)
1981 1982 1983 1984 1985 1986 1987 1988 1989 1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004
From Wikipedia: A vole is a small rodent resembling a mouse but with a stouter body, a shorter hairy tail, a slightly rounder head, and smaller ears and eyes. There are approximately 70 species of voles; they are sometimes known as meadow mice or field mice in America. The voles, together with the lemmings and the muskrats, form the subfamily Arvicolinae.
Species of voles can be found in Europe, Asia, North Africa and North America, and tundra areas.
Depending on the species, the vole's diet consists of seeds, tubers, conifers needles, bark, various green vegetation such as grass and clover, and insects.
Many carnivores such as wolves, owls, hawks, coyotes, foxes, weasels and cats ( i.e. Angus) eat voles. A common predator to voles is the short-eared owl.
The average life of a vole is 36 months. Voles rarely live longer than 12 months. The longest lifespan of a vole ever recorded was 18 months.
Reproductive Rate - predators breed at a linear rate. Voles breed at a geometric rate. Vole's population seems to be controlled by periodic crashes. Cause of crashes is not well known.
How many species of voles are there in the United States? How many species of microtus are there in the United States?
Animal Diversity Web University of Michigan Museum of Zoology
"Habitat - California voles inhabit areas of broad-leaved chaparral, oak woodlands, and grasslands along the Pacific Coast in northern Baja California to central Oregon. This species has a restricted distribution, which is possibly due to relic populations. It seems to utilize unusual habitats in California compared to other species of voles throughout the North American continent. Marshy ground, saltwater and freshwater locations, wet meadows, coastal wetlands and dry, grassy hillsides are the preferred macrohabitats of this species.
"Ovulation in M. californicus is induced by copulation. This species experiences a post-partum estrus, and breeding can occur within fifteen hours after young are born. This allows up to 4 or 5 litters a season"
Voles can breed every three weeks under good conditions.
The breeding season seems to heavily rely on the wet season in non-coastal parts of California. In coastal populations breeding is aseasonal.
Number of offspring is 1 to 11; average of 4 or 5.
Gestation period is 22 days.
Time of weaning is 2 weeks.
Time to independence is 2 weeks.
Age at sexual or reproductive maturity (female) is 21 days.
Age at sexual or reproductive maturity (male) is 25 to 42 days.
"There are a great number of vole predators, including coyotes, kestrels, hawks, weasels, kits, owls, snakes, herons, egrets, and feral cats. Because of their rapid reproduction and periody high population densities, these voles are a keystone prey species."
From Michael H. Ferkin: (http://biology.memphis.edu/ferkin.htm) "Microtene rodents are best known for their unique population dynamics that include spectacular increases in population density (more than 15,000 voles/hectare) followed by equally remarkable 'crashes' (less than 1 vole/hectare). Although the causes of microtene population fluctuations remain hidden, many researchers stress the importance of behavioral interactions as a mechanism that regulates vole demography. It has been hypothesized that seasonal plasticity of behavior and reproduction among individuals profoundly affect microtene social organization and demography. My approach to the study of animal behavior includes both proximate and ultimate levels of analysis."
Population ecology is the study of the reasons why the numbers of an animal species goes up and down. What are the factors that cause populations to fluctuate? Farley Mowet, in his book Never Cry Wolf, retells an Eskimo myth that explains the relationship between the wolf and the caribou; between predator and prey.
Initially the caribou lived in an environment without the wolves and the caribou thrived. They ate well and prospered. But soon their numbers were too great for their food supply. They started to starve and the population suffered as the weak caribou reproduced. So the wolf was brought into their land. The wolf preyed upon the weakest of the caribou. Eventually the caribou became strong again because the wolf takes out the weakest of the herd and only the strong survive and only the strong reproduce.
In a perfect situation each species of animal has an intrinsic rate of natural increase. This increase would continue until the population was at the point where it could be supported by the available resources. The small letter "r" is used to represent the rate of population increase and the letter "k" is used to represent the point where the population can be supported by the available resources.
Of course the relation between predator and prey is more complicated than the Eskimo story. There are fluctuations in the resources. There are fluctuations that occur to the prey items. There are fluctuations that occur to the predators. There are fluctuations that occur to the weather that affect the food that affects the prey items that affects the predators.
Studying the population ecology of the White-tailed Kite is a process of identifying the factors involved in a sustainable population. A sustainable population for one species is not necessarily sustainable for another species. It was realized after the Passenger Pigeon became extinct, that their natural history required that they maintain a very large population; a population so large that it came into conflict with the growing nation of America.
The White-tailed Kite is a microtene obligate. The main prey item for the White-tailed Kite is the vole; a microtine mammal. Part of the population ecology of the vole is that its population numbers are very erratic. In a good year they can have an incredible density of 15,000 voles per hectare. Within a year or two this density can shrink to one vole per hectare when the food and the water become scarce. The challenge for the White-tailed Kite is to be able to successfully respond to these changes.
Population ecology also includes the effect of one species on another species. This includes the human animal species. Many historical accounts talk of the ease which indiscriminate hunters were able to shoot the White-tailed Kite.
From Odum's Fundamentals of Ecology: p. 239
|
Prior to 1907 there were estimated to be about 4,000 deer and a good population of predators - pumas and wolves - on the Kaibab plateau, an area of about 700,000 acres on the north side of the Grand Canyon of Arizona. Between 1907 and 1923 a concerted effort was made to remove the predators. By 1925 the deer population had increased to 100,000 which was far beyond the carrying capacity of the vegetation. Everything in reach - grass, tree seedlings, shrubs - was eaten and the whole area gave the appearance of a huge over-grazed and over-browsed pasture. In two winters 40 per cent of the enormous herd starved to death, and decline continued to about 10,000. The range continues to be depleted, and damage to forest reproduction will be evident for a long time. It has been estimated that the original range could have supported no more than 30,000 deer. Thus, the predator-prey interaction was maintaining a relatively stable equilibrium, with the deep population being held well below the point where its own food supply would be depleted. |
One of the goals of population ecology is to understand any patterns that could exist in nature that would help explain the ecology of particular species.
|
To do science is to search for repeated patterns, not simply to accumulate facts, and to do the science of geographical ecology is to search for patterns of plant and animal life that can be put on a map. Robert MacArthur, Geographical Ecology |
Much evidence points to hunters being the cause of the kite's brush with extinction. Between 1880 and 1930 it seems very possible that an effort was made to exterminate all predators. The White-tailed Kite was an easy, convenient target given its white plumage, and its tendency to hover low over the fields.
There is a complicated history/relationship between ornithology and hunting. It is not the simple relationship that many people might expect. It is necessary, initially, to indicate that the discussion about hunting includes only legal hunting. There are people who do not purchase licenses, and there are people who kill animals that they know they shouldn't kill. They are an embarrassment to the hunting community at large.
There are also birdwatchers that hate all hunters and consider the whole process of hunting repellent. But they are eager to go on field trips to refuges that are bought and maintained by the fees paid by hunters. In California much of the land that sustains the large wintering populations of ducks, geese, other waterbirds, and hawks, is paid for from the fees collected from hunters. It has been suggested that hunters have bought a lot more strategic land than have birdwatchers.
This is not to say that there have not been egregious examples of ruthless, damaging hunting. In the 19th century the market hunters devised methods of slaughtering Passenger Pigeons so that they killed an estimated 10 billion birds in a period of a few years. It is miraculous that many species of shorebirds did not also become extinct since they were also the target of market hunters.
Additionally, there was a period of time when people had a mistaken notion of community ecology. People determined that predators were bad animals to have around because they killed songbirds, livestock, pets, etc., and because of this they should be destroyed.
One of the classic studies of the effect of the effort to kill all predators took place in Kern County, California around 1927. The whole community organized to kill as many predators as possible. So many predators were killed so quickly that the rodent population skyrocketed and that had a very dire effect on all aspects of life. At one point there were so many rodents on the roads being run over by cars, that cars were having problems staying on the roads because the roads became slippery due to the number of dead rodents.
Hunting still represents the wild-west independent nature of the United States. There is a culture of hunting that is not very old. In the life history accounts found in A. C. Bent's Life Histories of North American Birds it is possible to find candid comments by hunters who are the ornithologists and comments from them about the hunters who are not ornithologists.
There was still the concept that the resources were infinite. They did not have to worry about how many animals were shot because there would always be replacement resources. Extinction was not understood. The Passenger Pigeon and the Carolina Parakeet were exceptions not the rule.
Between the Kaibab Peninsula's deer population and Kern County anti-predator campaign it seems that the citizens did not understand the prey/predator relationship so classically outlined in the Eskimo myth.
The purpose of the lesson plans is to provide students an opportunity to do real research about a current situation.
These lesson plans are dynamic because the questions that they ask are mostly research challenges as opposed to figuring out right and wrong answers.
These lesson plans are also unique because they utilize the Internet to provide data to work with; data that previously would have taken hours to gather.
One of the goals is to utilize life sciences and student initiative to do this research.
The study of birds has consistently been regarded as a hobby but not a science. Ian Newton in his book Population Ecology of Raptors notes that the life histories of birds of prey are seldom used as examples in the studies of prey/predator relations and population ecology in general. The study of a population of animals is a scientific study.
Examination of a predator whose main niche is to prey upon rodents whose numbers can explode and cause health problems to the community is a serious consideration.
Work done by students can be sent to birdcentral.net and posted.
Caution: Between the years 1960 and 1980 the White-tailed Kite had its name changed to the Black-shouldered Kite. If you try to gather data from the Audubon CBC database for any portion of years between 1960 and 1980 you will need to use the name Black-shouldered Kite instead of White-tailed Kite.
|
|
||
|
12. Microtene Bust |
1. How many Voles?
|
Lesson Plan Name |
How many Voles? |
|
Subject Area |
Zoology |
|
Topic Area |
Life Science |
|
Grade Level |
6 -12 |
|
Objectives |
To use mathematics to help determine food web parameters |
|
Standards |
|
|
Materials |
Calculator |
|
Instructional Procedure |
During the months between October and March much of California is inhabited by hawks that have left the far north to be in a more temperate climate and more importantly, to be where there are more prey animals available. What you need to do is to use the number of predatory birds to determine an estimate on how many California Voles there might be in the area. If X number of birds spend 180 days in a definable territory then we can assume that they are finding food to keep them alive. |
|
Activities |
Select a Christmas Bird Count (CBC) that has a good diversity of birds that share the same diet as the White-tailed Kite. A table (Appendix H) has been provided that identifies a group of birds that share similar diets. For this study we have determined that the ready-to-eat vole weighs 25 grams. Pick a CBC that has a number of vole predators. Set up a formula and determine how many voles theoretically would be eaten to support this number of birds for a winter period. |
|
Resources |
Use Appendix H for energetic requirements of selected hawks and owls. |
|
Additional Questions |
1. What other species in the CBC eats Microtus species? 2. Dunk reports that there needs to be a certain availability of voles to make hunting successful. He suggests 1500 voles per territory and that an average territory is 10 hectares. 3. Obviously this is a pretty narrow scenario. Many variables are not included. List some of the major variables that would need to be considered to make this a more accurate study of wintering food needs. 4. Take ten similar counts and run the same equation. Are there any similarities in the numbers? |
|
Lesson Plan Name |
The size of the territory |
|
Subject Area |
Zoology |
|
Topic Area |
Life Science |
|
Grade Level |
6 -12 |
|
Objectives |
To learn more about how different species utilize the land. To understand the logistics of a life cycle. |
|
Standards |
|
|
Materials |
Calculator |
|
Instructional Procedure |
How much territory does a kite need to breed? If there are 140 birds, 70 pairs, in 150 square miles, how many of those pairs can breed? What are the factors that are involved with this question? |
|
Activities |
1. From the Appendix A pick a count that has an appropriate number of kites. 2. For this question we will assume that the number of wintering kites will be the number of breeding kites. 3. Translate the CBC area into hectares (See Appendix G) 4. Given a breeding area of 10 hectares how many breeding territories could there be in this area? |
|
Resources |
|
Lesson Plan Name |
Develop a Scenario |
|
Subject Area |
Science |
|
Topic Area |
Life Sciences |
|
Grade Level |
8 - 12 |
|
Objectives |
To interpret the meaning of a bird census. |
|
Standards |
|
|
Materials |
|
|
Instructional Procedure |
Here is an opportunity to become a reporter/mystery writer, manager. The CBCs are filled with many numbers but what those numbers tell us is up to us to find out. The challenge here is to take a series of counts and develop an explanation of what you think the numbers mean. How you do this is very much up to you. You may compare a number of different counts. You may want to take a ten year period of one count that shows a wide range of numbers. Your job is to develop a scenario for what you pick that helps to explain what the numbers mean. |
|
Activities |
Identify a sub-set of counts. Graph the numbers that you use and develop a scenario that suggests what the graphs might mean. Find some outside resources that pertain to your set of numbers that helps to explain them more fully. |
|
Resources |
|
Lesson Plan Name |
Average, Mean, median |
|
Subject Area |
Mathematics/Science |
|
Topic Area |
Life Science |
|
Grade Level |
8 -12 |
|
Objectives |
To connect statistical analysis with bird census data |
|
Standards |
|
|
Materials |
Calculator |
|
Instructional Procedure |
The concepts of average, mean, and mode are used to express different aspects of a group of numbers. Through the use of these terms we can better understand the character of the data that has been gathered. |
|
Activities |
1. Select a CBC that has at least 20 different counts. 2. Determine the average, the mean, and the mode for each count. 3. Discuss what these three expressions tell you about the character of these counts. 4. Which one of the three terms is more appropriate for this count and why. |
|
Resources |
|
Lesson Plan Name |
Graphing |
|
Subject Area |
Mathematics/Science |
|
Topic Area |
|
|
Grade Level |
6 - 12 |
|
Objectives |
To introduce the concept that zoological results can be graphed |
|
Standards |
|
|
Materials |
Graphing software |
|
Instructional Procedure |
Graphs provide an opportunity to visually identify the nature of a set of numbers. We can look at a series of numbers and get a sense of what is known but the numbers along with a graph can quite often show trends in the numbers more dramatically. |
|
Activities |
1. Pick a CBC with at least 20 years of results. (See Appendix A) 2. Graph the results and be sure to label the axes. 3. Interpret what the graph visually shows. |
|
Resources |
|
Lesson Plan Name |
Population Dynamics of the California Vole |
|
Subject Area |
Zoology |
|
Topic Area |
Life Sciences |
|
Grade Level |
Grades 5 - 7 |
|
Objectives |
To better understand the logistics of animal populations. To better understand the prey/predator relationship. |
|
Standards |
|
|
Materials |
A calculator |
|
Instructional Procedure |
The Prey-Predator relationship is sometimes a difficult concept for someone to understand who has been told to protect animals. But to understand the place of raptors in an environment we need to consider what would happen if they weren't present. By considering the ability of mice to reproduce with a quick turn around very dramatically communicates the necessity of predators. |
|
Activities |
The California Vole has a gestation period of 3 weeks. They have, on the average 6 to 10 babies. Those babies mature in about 5 weeks. That means they are now ready to have their own babies. Anticipate that there will be an equal percent of 50% male babies and 50% female babies. If you start on January 1 with a pair of voles and there are no predators or disease, approximately how many mice will you end up with by June 30; December 31? If you start with a pair of voles on Jan. 1 theoretically when would you have 1500 voles? |
|
Resources |
Some birds who are very responsible for killing voles are: White tailed Kite, Barn Owl, Gt. Blue Heron, Common Crow. |
|
Lesson Plan Name |
|
|
Subject Area |
Zoology |
|
Topic Area |
Life Sciences/Math |
|
Grade Level |
Grades 3 - 6 |
|
Objectives |
To understand the mathematical description of prey-predator relations. To understand the difference between linear and geometric growth. |
|
Standards |
|
|
Materials |
scratch paper |
|
Instructional Procedure |
This is another exercise that expresses how populations can grow when there aren't balancing factors. Students are offered the choice between being paid a lump sum of $100,000 for one month of work or being paid a penny a day but having the amount doubled every day for the month. This exercise helps to demonstrate the difference between linear growth and geometric growth. |
|
Activities |
1. You are going to work for one month and your employer has given you the choice between getting paid $100,000 for the month or getting one penny a day. But that penny gets doubled every day for 31 days. So on day one you get .01 and on day two you get .02. The amount gets doubled every day for 31 days. It's your choice. 2. What does this have to do with the California Vole? |
|
Lesson Plan Name |
Profile of a CBC |
|
Subject Area |
Science |
|
Topic Area |
Life Science |
|
Grade Level |
9 - 12 |
|
Objectives |
To understand the concept of ecology. |
|
Standards |
|
|
Materials |
|
|
Instructional Procedure |
We know the basic details of a Christmas Bird Count. It is a bird census that takes place on a particular day during the last two weeks of each year. The participants count and list all the birds found over a 24 hour period. The count takes place in a circle with a diameter of 15 miles. Your profile will add to these details by including the other organisms involved from the human organisms to the domestic organisms to the wild non-bird organisms that don't get counted. |
|
Activities |
Pick one of the CBCs from the Audubon Historical Database that includes White-tailed Kites. 1. What industry is in the area? 2. What farms and ranches are currently working? How many farm animals? 3. Is there any hunting going on in the area on the same day? 4. How many domestic pets? 5. What has been the economic history of the area? |
|
Resources |
|
|
Lesson Plan Name |
History of California Agriculture |
|
Subject Area |
History |
|
Topic Area |
|
|
Grade Level |
6 - 12 |
|
Objectives |
To better understand the role of agriclture in the ecology of the state. |
|
Standards |
|
|
Materials |
|
|
Instructional Procedure |
Create a brief history of the major changes in California agriculture during the period from1880 to 1980. Number of farms Number of ranches Irrigation access |
|
Activities |
|
|
Resources |
|
Lesson Plan Name |
Sim City Meets the White-tailed Kite - Defining K |
|
Subject Area |
Science |
|
Topic Area |
Life Science |
|
Grade Level |
8 - 12 |
|
Objectives |
To better understand the dynamics of an animal community. |
|
Standards |
|
|
Materials |
|
|
Instructional Procedure |
|
|
Activities |
You have suddenly acquired a management opportunity. You get to create your own maximum White-tailed kite environment. What would an ideal kite environment look like? In a short essay develop an ideal White-tailed Kite environment. Take into consideration what resources will be needed on a year round basis. How many voles are there? How many other mammals? Where is the water coming from? What is the human involvement? How many wintering kites are there and how many breeding kites? |
|
Resources |
|
Lesson Plan Name |
The White-tailed Kite as an Indicator Species |
|
Subject Area |
Zoology |
|
Topic Area |
Life Science |
|
Grade Level |
9 - 12 |
|
Objectives |
To understand what an Indicator Species indicates. |
|
Standards |
|
|
Materials |
|
|
Instructional Procedure |
The White-tailed Kite has very specific geographical needs and prey requirements. It prefers fresh water grasslands below 2000 feet with plenty of California Voles running around. (About 1500 per hectare.) Given the number of CBC counts in California that have White-tailed Kites there is a wide variety of kite density that can be found in California. |
|
Activities |
Pick different counts that have a wide variety of kite density: 1, 10, 30, 50 80, 100 kites per count. Anticipate what the environment looks like given the requirement of the number of kites. If there are over 150 kites on the CBC that is prime kite environment. If there are less than 10 then we might expect a small remnant of grassland. |
|
Resources |
|
Lesson Plan Name |
Microtine Bust |
|
Subject Area |
Science |
|
Topic Area |
Life Science |
|
Grade Level |
9 - 12 |
|
Objectives |
To understand the differences in the behavior of a predator specialist compared to a predator generalist. |
|
Standards |
|
|
Materials |
|
|
Instructional Procedure |
Imagine that the California Vole population has bottomed out. In this scenario there are only two voles per hectare. Their population will rebound next year but right now the rodent predators have to improvise. |
|
Activities |
Write a short essay that discusses life without the voles. 1. Describe the strategies of the specialist White-tailed Kite and the generalist Red-tailed Hawk. 2. How does life as a generalist compare to life as a specialist? What happens to the White-tailed Kite if it leaves its preferred habitat? 3. What does it mean to be a nomadic hunter? |
A. White-tailed
Kite Numbers in Selected California Christmas Bird
Counts
White-tailed Kite
Numbers for the Benicia Christmas Bird Counts
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
White-tailed Kite
Numbers for the Sacramento National Wildlife Refuge
Christmas Bird Counts
White-tailed Kite
Numbers for the Los Banos Christmas Bird Counts
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
||||||||
|
|
|
|
White-tailed Kite
Numbers for the Folsom Christmas Bird Counts
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|||||||
|
|
|
|
|
White-tailed Kite
Numbers for the Santa Barbara Christmas Bird
Counts
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
White-tailed Kite
Numbers for the Palo Alto Christmas Bird Counts
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
||||||||||
|
|
White-tailed Kite
Numbers for the San Jose Christmas Bird Counts
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|||
|
|
|
|
|
|
|
|
|
White-tailed Kite
Numbers for the Ano Nuevo Christmas Bird Counts
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
White-tailed Kite
Numbers for the Peace Valley Christmas Bird
Counts
White-tailed Kite
Numbers for the Orange County (Coastal) Christmas Bird
Counts
B. Highcounts for
White-tailed Kites for CBC's between 1960 and 2004.
All locations are in California unless
otherwise indicated (1992, 1997). 110 204 137 202 Los Banos
C. Total Numbers
for White-tailed Kite (Black shouldered Kite) for 1960 -
2004 by state Year 1960 1961 1962 1963 1964 1965 1966 1967 1968 1969 1970 1971 1972 1973 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988 1989 1990 1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004
D. 1996 California Christmas Bird
Count with observations of at least one Northern
Harrier
|
|
|||
|
|
|
||
|
Land Area of State: |
|
Land Area of State: |
|
|
Cultivatable Land |
|
||
|
Land in 150,360 Farms |
|
||
|
Land Drained for Agriculture |
|
||
|
Potentially irrigable land |
|
||
|
Area under irrigation |
|
||
|
Non-agricultural land |
|
||
|
Original forested area |
|
||
|
Present forested area |
|
||
|
Present virgin timber |
|
||
|
20 National forests |
|
||
|
Grazing land in forests |
|
||
|
Public domain |
|
||
|
People in California |
|
|
|
|
Urban (73.3%) |
|
||
|
Rural |
|
||
|
Farm |
|
||
|
Number per square mile |
|
|
|
|
Per cent increase in 10 years |
|
|
|
|
Hunters' licenses (1935) |
|
|
|
Ano Nuevo Los Banos Peace Valley Auburn Marin Pt. Reyes Benicia Moss Landing Santa Barbara Centerville Beach Moro Bay Santa Maria Contra Costa Orange County (coastal) San Jacinto Lake Folsom Orange County Sacramento Hayward Oceanside Stockton Hollister Palo Alto Thousand Oaks
640 Acres to 1 Square mile Hectare equals 2.471 acres Area of a circle, A = pi
(r2) pi = 3.14 (approximately) Diameter of a CBC is 15
miles. The radius (r) of a circle is half the
diameter. hectare = 100 M x 100 M
H. Selected Hawks and Owls of
California that eat over 50% Voles
White-tailed Kite Red-tailed Hawk Red-shouldered Hawk Rough-legged Hawk Ferruginous Hawk Northern Harrier (Marsh
Hawk) American Kestral Barn Owl Short-earred Owl Long-earred Owl
American Ornithologists Union. 1983. Checklist of North American Birds - Sixth Edition. American Ornithologists' Union, Washington, D.C..
Bent, A. C. Life Histories of North American Birds of Prey, V. 1. Dover Publications. 1961
Clark, S. W. and B. K. Wheeler. Peterson Field Guides - Hawks Houghton Mifflin Company1987
CPIF (California Partners in Flight). 2000. Version 1.0. The draft grassland bird conservation plan: a strategy for protecting and managing grassland habitats and associated birds in California (B. Allen, lead author). Point Reyes Bird Observatory, Stinson Beach, Ca. http://www.prbo.org/CPIF/Consplan.html.
Dawson, W. L. The Birds of California, Volume 3. South Moulton Company, 1923.
del Hoyo, J., A. Elliot & J. Sargatal (Eds.) Handbook of the Birds of the World, Volume 2. Lynx Edicions. Barcelona 1994
Dunk, J. R. 1995. White-tailed Kite (Elanus leucurus). In The Birds of North America, No. 178 (A. Poole and F. Gill, eds.). The Academy of Natural Sciences, Philadelphia, and The American Ornithologists' Union, Washington, D.C.
Dunk, J. R., & R. J. Cooper. Territory-size Regulation in Black-shouldered Kites. Auk, Vol. 111. No. 3 July 1994
Ehrlich, P. R., D. S. Dobkin, & D. Wheye. The Birder's Handbook. New York. Fireside Book. 1988.
Ellis, D. H. & G. Monson. White-tailed Kite Records for Arizona. Western Birds, Vol. 10, No. 3, 1979.
Garbrielson, I. N. & S. G. Jewett. Birds of the Pacific Northwest. Dover Publications, New York. 1970
Gill, R. Jr. Breeding Avifauna of the South San Francisco Bay Estuary. Western Birds, Vol. 8, No. 1, 1977
Gilliard, T. E. Living Birds of the World. Garden City. Doubleday and Company. 1958
Greenberg, R. & S. Droege. On the Decline of the Rusty Blackbird and the Use of Ornithological Literature to Document Long-term Population Trends. Conservation Biology, Vol. 13, No. 3, June 1999; pgs. 553 - 559.
Grinnell, J. A Distributional List of the Birds of California. Cooper Ornithological Club. 1915
Grinnell, J & A. H. Miller. The Distribution of the Birds of California. Cooper Ornithological Club. 1944
Hamerstrom, F. Harrier - Hawk of the Marshes. Washington, D.C.. Smithsonian Institution Press. 1986.
Henny, C. J. & J. T. Annear. A White-tailed Kite Breeding Record for Oregon. Western Birds, Vol. 9, No.3, 1978 p. 131-132.
Hobbs, R. J. & H. A. Mooney. Broadening the Extinction Debate: Population Deletions and Additions in California and Western Australia. Conservation Biology, Vol. 12, No. 2, April 1998.
Hoffmann, R. Birds of the Pacific States. Houghton Mifflin Company. 1927.
Koenig, W. D. Spatial Autocorrelation in California Land Birds. Conservation Biology, Vol. 12, No. 3, June 1998; pgs. 612 - 619.
Koford, R. R., J. B. Dunning Jr., C. A. Ribic, & D. M. Finch. A Glossary for Avian Conservation Biology. Wilson Bulletin, 106 (1), 1994. pp. 121 - 137.
|
|
|
Le Boeuf, B., and S. Kaza. The Natural History of Ano Nuevo. Pacific Grove. Boxwood Press, 1981
Linsdale, J. M. Preservation of Birds in California. Condor Vol. 39
MacArthur, R. H. Geographical Ecology - Patterns in the Distribution of Species. Princeton. Princeton University Press. 1972.
MacWhirter, R. B., and K. L. Bildstein. 1996. Northern Harrier (Circus cyaneus). In The Birds of North America, No. 210 (A. Poole and F. Gill, eds.). The Academy of Natural Sciences, Philadelphia, and The American Ornithologists' Union, Washington, D.C.
Moore, J. White-tailed Kite. http://www.prbo.org/calpif/htmldocs/species/grassland/wtkiacct.html Downloaded on 2/17/2005
Mowet, F. Never Cry Wolf
Newton, I. Population Ecology of Raptors Vermillion, North Dakota. Buteo Books, 1979.
Newton, I. Population Limitation in Birds. San Diego. Academic Press, 1998
Odum, E. P. Fundamentals of Ecology (Second Edition). Philadelphia. W. B. Saunders Company, 1959
Peronne, L. 2002. "Microtus californicus" (On-line), Animal Diversity Web. Accessed December 07, 2007 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Microtus_californicus.html
Peterson, R. T. A Field Guide to Western Birds. Houghton Mifflin Company, 1960
Pickwell, G. Requiem for the White-tailed Kites of Santa Clara County. Condor. 1931
Pickwell, G. The White-tailed Kite. Condor Volume 32, Number 5, pp. 221 - 239.
Price, J., S. Droege & A. Price. The Summer Atlas of North American Birds. New York. Academic Press, 1995
Roberson, D. & C. Tenney. Atlas of the Breeding Birds of Monterey County California. Monterey Peninsula Audubon Society. 1993
Root, T. Atlas of Wintering North American Birds. Chicago. University of Chicago Press. 1988.
Scott, M. J. Preserving and restoring avian diversity: A search for solutions. In A Century of Avifaunal Change in Western North America. Studies in Avian Biology No. 15. Cooper Ornithological Society, 1994.
Shuford, W. D. The Marin County Breeding Bird Atlas. Bushtit Books. 1993.
Small, A. The Birds of California. Winchester Press, New York. 1974.
Small, A. California Birds - Their Status and Distribution. Vista. Ibis Publishing Company. 1994
Warner, J. S. & R. L. Rudd. Hunting by the White-tailed Kite (Elanus Leucurus). The Condor 1975
White, C. M. Population Trends in the landbirds of western North America. in A Century of Avifaunal Change in Western North America. Studies in Avian Biology No. 15. Cooper Ornithological Society, 1994
|
|
|
|